They multiplied and filled the oceans and seas with oxygen encouraged the later developm

问题补充：

They multiplied and filled the oceans and seas with oxygen, encouraged the later development of early shellfish and all sorts of fish.

A. that

B. where

C. when

D. which

答案：

A. that

B. where

C. when

(答案→)D. which

解析：可联系答.案.网客服索取。